Born in Aberystwyth, Wales, he started his career with the Royal Shakespeare Company. He created the title role in the original production of the play The Life and Adventures of Nicholas Nickleby, winning both an Olivier Award and a Tony Award for Best Actor in a Play in 1982 for it. He also starred in the original production of The Real Thing by Tom Stoppard in London in 1984. Rees became an American citizen in 1989, and in the 1990s, continued his work in the theatre, both as an actor and a director.
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